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Reply to "Can a R4LC replace an R2LC on this Engine"

The ERR R4LC is 100% plug compatible with the Lionel R2LC-C08.  However, note that the ERR R4LC is NOT compatible with any Legacy coded R4LC, those would be the -S1 thru -S5 versions.  For  Legacy locomotive, you really need a Legacy code R4LC.

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