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Reply to "Electrocouplers"

I was converting a Williams PA 1 A-A set to R4LC command with an ERR Cruise Commander, and when putting in a couple Lionel electro-couplers the question came to mind while creating the link to the coupler added to the dummy unit.  "Why not just run a one wire tether," I thought?  (I ran both.)  The answers seem to indicate that depending on the circumstances a board may use AC or DC to operate a coupler.  If DC, then a DC ground is needed which is found in the board.  So, if correct, why are there two wires to the motherboard with AC, which the R4LC situation apparently is?  Aw heck, I don't even know enough to know what the reference to "reference" means?  Good grief.    

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