I was converting a Williams PA 1 A-A set to R4LC command with an ERR Cruise Commander, and when putting in a couple Lionel electro-couplers the question came to mind while creating the link to the coupler added to the dummy unit. "Why not just run a one wire tether," I thought? (I ran both.) The answers seem to indicate that depending on the circumstances a board may use AC or DC to operate a coupler. If DC, then a DC ground is needed which is found in the board. So, if correct, why are there two wires to the motherboard with AC, which the R4LC situation apparently is? Aw heck, I don't even know enough to know what the reference to "reference" means? Good grief.
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